Strict consensus of six most parsimonious trees showing characters (above) and state (below). Bootstrap support values are given in parentheses (tree length=275 steps, ensemble consistency index (CI) = 0.39, ensemble retention index (RI) = 0.64). C. archratiras is referred to as “Orapa fossil”. Character set from Bai et al. (2011).

  Part of: Woolley C (2016) The first scarabaeid beetle (Coleoptera, Scarabaeidae, Melolonthinae) described from the Mesozoic (Late-Cretaceous) of Africa. African Invertebrates 57(1): 53-66.